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Confirmed that DNA is genetic material through experiments with a virus called bacteriophage.

Experiments showed that DNA is the transforming principle not protein.

Influenced by data from other researchers to create a model of metal and wood to figure out the structure of DNA is a double helix twisted ladder

Concluded that some material must be transferred from heat-killed S bacteria to the live R bacteria. Called this material the "Transforming principle"

Used X-ray crystallography showed photographs of DNA in an X surrounded by a circle indicating it was a helix.

Analyzed DNA of several organisms and found that the same four bases are found in the DNA of all organisms. Found that the amount of A = T and C = G.